About the Authors

Hai-Lun He

Contributed equally to this work with: Hai-Lun He, Jun Guo

Affiliation State Key Laboratory of Microbial Technology, Marine Biotechnology Research Center, Shandong University, Jinan, China

Jun Guo

Contributed equally to this work with: Hai-Lun He, Jun Guo

Affiliation State Key Laboratory of Microbial Technology, Marine Biotechnology Research Center, Shandong University, Jinan, China

Xiu-Lan Chen

Affiliation State Key Laboratory of Microbial Technology, Marine Biotechnology Research Center, Shandong University, Jinan, China

Bin-Bin Xie

Affiliation State Key Laboratory of Microbial Technology, Marine Biotechnology Research Center, Shandong University, Jinan, China

Xi-Ying Zhang

Affiliation State Key Laboratory of Microbial Technology, Marine Biotechnology Research Center, Shandong University, Jinan, China

Yong Yu

Affiliation SOA Key Laboratory for Polar Science, Polar Research Institute of China, Shanghai, China

Bo Chen

Affiliation SOA Key Laboratory for Polar Science, Polar Research Institute of China, Shanghai, China

Bai-Cheng Zhou

Affiliation State Key Laboratory of Microbial Technology, Marine Biotechnology Research Center, Shandong University, Jinan, China

Yu-Zhong Zhang

zhangyz@sdu.edu.cn

Affiliation State Key Laboratory of Microbial Technology, Marine Biotechnology Research Center, Shandong University, Jinan, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HLH XLC YZZ. Performed the experiments: HLH JG. Analyzed the data: HLH JG BBX. Contributed reagents/materials/analysis tools: XYZ YY BC BCZ. Wrote the paper: HLH JG XLC YZZ.