About the Authors

Ke Yuan

Affiliation Department of Biology, College of Science and Technology, Temple University, Philadelphia, Pennsylvania, United States of America

Zhaorui Lian

Affiliation Department of Biology, College of Science and Technology, Temple University, Philadelphia, Pennsylvania, United States of America

Bill Sun

Affiliation Department of Biology, College of Science and Technology, Temple University, Philadelphia, Pennsylvania, United States of America

Marcia M. Clayton

Affiliation Department of Biology, College of Science and Technology, Temple University, Philadelphia, Pennsylvania, United States of America

Irene O. L. Ng

Affiliation Department of Pathology, University of Hong Kong, Queen Mary Hospital, Pokfulam, Hong Kong

Mark A. Feitelson

feitelso@temple.edu

Affiliations Department of Biology, College of Science and Technology, Temple University, Philadelphia, Pennsylvania, United States of America, Center for Biotechnology, Sbarro Health Research Organization, College of Science and Technology, Temple University, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KY ZRL BS MMC IOLN MF. Performed the experiments: KY ZRL BS MMC IOLN MF. Analyzed the data: KY ZRL BS MMC IOLN MF. Contributed reagents/materials/analysis tools: KY ZRL BS MMC IOLN MF. Wrote the paper: KY ZRL BS MMC IOLN MF.