About the Authors

Guang-Jie Zhu

Contributed equally to this work with: Guang-Jie Zhu, Fang Wang

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Fang Wang

Contributed equally to this work with: Guang-Jie Zhu, Fang Wang

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Chen Chen

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Lin Xu

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Wen-Cheng Zhang

Affiliations MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China, Zhejiang Provincial Key Lab for Technology & Application of Model Organisms, School of Life Sciences, Wenzhou Medical College, University Park, Wenzhou, China

Chi Fan

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Ya-Jing Peng

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Jie Chen

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Wei-Qi He

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Shi-Ying Guo

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Jian Zuo

Affiliation Department of Developmental Neurobiology, St. Jude Children's Research Hospital, Memphis, Tennessee, United States of America

Xia Gao

xiagao213@yahoo.com.cn (XG); zhums@nju.edu.cn (M-SZ)

Affiliation MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China

Min-Sheng Zhu

xiagao213@yahoo.com.cn (XG); zhums@nju.edu.cn (M-SZ)

Affiliations MOE Key Laboratory for Model Animal and Diseases Studies, Nanjing Drum Tower Hospital and Model Animal Research Center of Nanjing University, Nanjing, China, Zhejiang Provincial Key Lab for Technology & Application of Model Organisms, School of Life Sciences, Wenzhou Medical College, University Park, Wenzhou, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: G-JZ FW JZ XG M-SZ. Performed the experiments: G-JZ FW LX W-CZ CF Y-JP JC W-QH CC S-YG M-SZ. Analyzed the data: FW CC LX. Contributed reagents/materials/analysis tools: G-JZ FW JZ. Wrote the paper: G-JZ XG M-SZ.