About the Authors

Kyle K. Pitchers

Affiliations Departments of Anatomy and Cell Biology, Schulich School of Medicine and Dentistry, The University of Western Ontario, London, Ontario, Canada, Departments of Molecular and Integrative Physiology, University of Michigan, Ann Arbor, Michigan, United States of America

Susanne Schmid

Affiliation Departments of Anatomy and Cell Biology, Schulich School of Medicine and Dentistry, The University of Western Ontario, London, Ontario, Canada

Andrea R. Di Sebastiano

Affiliations Departments of Anatomy and Cell Biology, Schulich School of Medicine and Dentistry, The University of Western Ontario, London, Ontario, Canada, Departments of Molecular and Integrative Physiology, University of Michigan, Ann Arbor, Michigan, United States of America

Xu Wang

Affiliation Departments of Anatomy and Cell Biology, Schulich School of Medicine and Dentistry, The University of Western Ontario, London, Ontario, Canada

Steven R. Laviolette

Affiliation Departments of Anatomy and Cell Biology, Schulich School of Medicine and Dentistry, The University of Western Ontario, London, Ontario, Canada

Michael N. Lehman

Affiliations Departments of Anatomy and Cell Biology, Schulich School of Medicine and Dentistry, The University of Western Ontario, London, Ontario, Canada, Departments of Molecular and Integrative Physiology, University of Michigan, Ann Arbor, Michigan, United States of America

Lique M. Coolen

coolenlm@umich.edu

Affiliations Departments of Anatomy and Cell Biology, Schulich School of Medicine and Dentistry, The University of Western Ontario, London, Ontario, Canada, Departments of Molecular and Integrative Physiology, University of Michigan, Ann Arbor, Michigan, United States of America, Department of Psychology, University of Michigan, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KP SS AD ML LC. Performed the experiments: KP SS AD XW. Analyzed the data: KP SS AD XW. Contributed reagents/materials/analysis tools: SS SL ML LC. Wrote the paper: KP LC.