About the Authors

Yan Han

Contributed equally to this work with: Yan Han, Guangyu Hou, Wenming Jiang

Affiliations China Animal Health and Epidemiology Center, Qingdao, China, College of Animal Science and Veterinary Medicine, Qingdao Agricultural University, Qingdao, China

Guangyu Hou

Contributed equally to this work with: Yan Han, Guangyu Hou, Wenming Jiang

Affiliation China Animal Health and Epidemiology Center, Qingdao, China

Wenming Jiang

Contributed equally to this work with: Yan Han, Guangyu Hou, Wenming Jiang

Affiliation China Animal Health and Epidemiology Center, Qingdao, China

Chunhua Han

Affiliation Institute of Animal and Husbandry Medicine, Beijing Academy of Agriculture and Forestry Sciences, Beijing, China

Shuo Liu

Affiliation China Animal Health and Epidemiology Center, Qingdao, China

Jie Chen

Affiliation China Animal Health and Epidemiology Center, Qingdao, China

Jinping Li

Affiliation China Animal Health and Epidemiology Center, Qingdao, China

Peng Zhang

Affiliations China Animal Health and Epidemiology Center, Qingdao, China, College of Animal Science and Veterinary Medicine, Qingdao Agricultural University, Qingdao, China

Baoxu Huang

Affiliation China Animal Health and Epidemiology Center, Qingdao, China

Yuehuan Liu

Affiliation Institute of Animal and Husbandry Medicine, Beijing Academy of Agriculture and Forestry Sciences, Beijing, China

Jiming Chen

jmchen66@yahoo.cn

Affiliation China Animal Health and Epidemiology Center, Qingdao, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: Jiming Chen. Performed the experiments: YH GH WJ CH SL Jie Chen JL PZ BH YL Jiming Chen. Analyzed the data: Jiming Chen YH. Contributed reagents/materials/analysis tools: Jiming Chen WJ YL. Wrote the paper: Jiming Chen.