About the Authors

Ziya Kalay

zkalay@icems.kyoto-u.ac.jp (ZK); akusumi@frontier.kyoto-u.ac.jp (AK)

Affiliation Institute for Integrated Cell-Material Sciences (WPI-iCeMS), Kyoto University, Kyoto, Japan

Takahiro K. Fujiwara

Affiliation Institute for Integrated Cell-Material Sciences (WPI-iCeMS), Kyoto University, Kyoto, Japan

Akihiro Kusumi

zkalay@icems.kyoto-u.ac.jp (ZK); akusumi@frontier.kyoto-u.ac.jp (AK)

Affiliations Institute for Integrated Cell-Material Sciences (WPI-iCeMS), Kyoto University, Kyoto, Japan, Institute for Frontier Medical Sciences, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TKF AK. Analyzed the data: ZK TKF AK. Wrote the paper: ZK TKF AK. Performed the simulations and developed the theory: ZK.