About the Authors

Wenke Zhou

Affiliation Department of Cell Biology and Physiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Juan Huang

Current address: National Institute of Biological Sciences, ZGC Life Sciences Park, Beijing, People's Republic of China

Affiliation Department of Cell Biology and Physiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Annie M. Watson

Affiliation Department of Cell Biology and Physiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Yang Hong

yhong@pitt.edu

Affiliation Department of Cell Biology and Physiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YH WZ JH. Performed the experiments: WZ JH AMW YH. Analyzed the data: WZ YH JH. Contributed reagents/materials/analysis tools: WZ JH AMW YH. Wrote the paper: YH WZ.