About the Authors

Brian Dimitroff

Affiliations Department of Genetics, Cell Biology and Development, Developmental Biology Center, University of Minnesota, Minneapolis, Minnesota, United States of America, Department of Biochemistry and Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Katie Howe

Contributed equally to this work with: Katie Howe, Adrienne Watson, Bridget Campion

Affiliation Graduate Program in Neuroscience, University of Minnesota, Minneapolis, Minnesota, United States of America

Adrienne Watson

Contributed equally to this work with: Katie Howe, Adrienne Watson, Bridget Campion

Affiliation Department of Genetics, Cell Biology and Development, Developmental Biology Center, University of Minnesota, Minneapolis, Minnesota, United States of America

Bridget Campion

Contributed equally to this work with: Katie Howe, Adrienne Watson, Bridget Campion

Affiliation Graduate Program in Neuroscience, University of Minnesota, Minneapolis, Minnesota, United States of America

Hyun-Gwan Lee

Affiliation Department of Biochemistry and Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Na Zhao

Affiliation Department of Biochemistry and Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Michael B. O'Connor

Affiliation Department of Genetics, Cell Biology and Development, Developmental Biology Center, University of Minnesota, Minneapolis, Minnesota, United States of America

Thomas P. Neufeld

Affiliation Department of Genetics, Cell Biology and Development, Developmental Biology Center, University of Minnesota, Minneapolis, Minnesota, United States of America

Scott B. Selleck

sbs24@psu.edu

Affiliations Department of Genetics, Cell Biology and Development, Developmental Biology Center, University of Minnesota, Minneapolis, Minnesota, United States of America, Graduate Program in Neuroscience, University of Minnesota, Minneapolis, Minnesota, United States of America, Department of Biochemistry and Molecular Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BDD KH AW HGL NZ BC SBS. Performed the experiments: BDD KH AW HGL NZ BC. Analyzed the data: BDD KH AW HGL NZ BC. Contributed reagents/materials/analysis tools: MBO TPN SBS. Wrote the paper: BDD HGL SBS.