About the Authors

Shin-ya Okamoto

Affiliation Department of Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, Hongo, Tokyo, Japan

Masamitsu Sato

masasato@biochem.s.u-tokyo.ac.jp

Affiliations Department of Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, Hongo, Tokyo, Japan, PRESTO, Japan Science and Technology Agency, Honcho Kawaguchi, Saitama, Japan

Takashi Toda

Affiliation Laboratory of Cell Regulation, Cancer Research UK, London Research Institute, Lincoln's Inn Fields Laboratories, London, United Kingdom

Masayuki Yamamoto

Affiliations Department of Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, Hongo, Tokyo, Japan, Kazusa DNA Research Institute, Kisarazu, Chiba, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SO MS MY. Performed the experiments: SO. Analyzed the data: SO. Contributed reagents/materials/analysis tools: TT MS. Wrote the paper: MS MY TT.