About the Authors

Keith B. Hengen

khengen@brandeis.edu

Affiliation Neuroscience Training Program, University of Wisconsin, Madison, Wisconsin, United States of America

Nathan R. Nelson

Affiliation Department of Comparative Biomedical Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Kyle M. Stang

Affiliation Department of Comparative Biomedical Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Stephen M. Johnson

Affiliations Neuroscience Training Program, University of Wisconsin, Madison, Wisconsin, United States of America, Department of Comparative Biomedical Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Stephanie M. Crader

Affiliation Department of Comparative Biomedical Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Jyoti J. Watters

Affiliation Department of Comparative Biomedical Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Gordon S. Mitchell

Affiliations Neuroscience Training Program, University of Wisconsin, Madison, Wisconsin, United States of America, Department of Comparative Biomedical Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Mary Behan

Affiliations Neuroscience Training Program, University of Wisconsin, Madison, Wisconsin, United States of America, Department of Comparative Biomedical Sciences, University of Wisconsin, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KH MB GM SJ JW. Performed the experiments: KH NN KS SC. Analyzed the data: KH. Contributed reagents/materials/analysis tools: MB SJ JW. Wrote the paper: KH.