About the Authors

Emma M. Baxter

Emma.Baxter@sac.ac.uk

Affiliation Animal Behaviour and Welfare, Scottish Agricultural College, Edinburgh, United Kingdom

Susan Jarvis

Affiliation Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Javier Palarea-Albaladejo

Affiliation Biomathematics and Statistics Scotland, Edinburgh, United Kingdom

Sandra A. Edwards

Affiliation School of Agriculture, Food and Rural Development, Newcastle University, Newcastle upon Tyne, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EB. Performed the experiments: EB. Analyzed the data: EB JP-A. Wrote the paper: EB. Contributted to sourcing funding: SJ SE. Contributed to editting manuscript: SJ SE. Contributed to statistical analysis: JP-A.