About the Authors

Shuai Wang

Contributed equally to this work with: Shuai Wang, An-Jing Chen, Li-Jie Shi

Affiliations The Key Laboratory of Plant Cell Engineering and Germplasm Innovation of Ministry of Education, School of Life Sciences, Shandong University, Jinan, Shandong, People's Republic of China, Wuhan Institute of Virology, Chinese Academy of Science, Wuchang, Hubei, People's Republic of China

An-Jing Chen

Contributed equally to this work with: Shuai Wang, An-Jing Chen, Li-Jie Shi

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation of Ministry of Education, School of Life Sciences, Shandong University, Jinan, Shandong, People's Republic of China

Li-Jie Shi

Contributed equally to this work with: Shuai Wang, An-Jing Chen, Li-Jie Shi

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation of Ministry of Education, School of Life Sciences, Shandong University, Jinan, Shandong, People's Republic of China

Xiao-Fan Zhao

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation of Ministry of Education, School of Life Sciences, Shandong University, Jinan, Shandong, People's Republic of China

Jin-Xing Wang

jxwang@sdu.edu.cn

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation of Ministry of Education, School of Life Sciences, Shandong University, Jinan, Shandong, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JXW SW XFZ. Performed the experiments: SW AJC LJS. Analyzed the data: SW JXW. Contributed reagents/materials/analysis tools: JXW XFZ. Wrote the paper: SW JXW. Performed the experiments and writing: SW.