About the Authors

Baoping Ren

Contributed equally to this work with: Baoping Ren, Dayong Li

Affiliation Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Dayong Li

Contributed equally to this work with: Baoping Ren, Dayong Li

Affiliations Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, College of Life Sciences, China West Normal University, Nanchong, Sichuan, China

Paul A. Garber

lim@ioz.ac.cn (ML); p-garber@illinois.edu (PAG)

Affiliation Department of Anthropology Program in Ecology and Evolutionary Biology, University of Illinois, Urbana, Illinois, United States of America

Ming Li

lim@ioz.ac.cn (ML); p-garber@illinois.edu (PAG)

Affiliation Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the study: BPR ML. Data collection and behavior observation: BPR DYL. Analyzed the data: BPR. Wrote the paper: BPR PAG ML.