About the Authors

Shuhei Nakamura

Affiliation Laboratory of Molecular Genetics for Reproduction, National Institute for Basic Biology, Okazaki, Japan

Ikuko Watakabe

Affiliation Laboratory of Molecular Genetics for Reproduction, National Institute for Basic Biology, Okazaki, Japan

Toshiya Nishimura

Affiliations Laboratory of Molecular Genetics for Reproduction, National Institute for Basic Biology, Okazaki, Japan, Department of Basic Biology, The Graduate University for Advanced Studies (SOKENDAI), Okazaki, Japan

Atsushi Toyoda

Affiliation Comparative Genomics Laboratory, National Institute of Genetics, Mishima, Japan

Yoshihito Taniguchi

Affiliation Department of Preventive Medicine and Public Health, School of Medicine, Keio University Shinanomachi 35, Tokyo, Japan

Minoru Tanaka

mtanaka@nibb.ac.jp

Affiliations Laboratory of Molecular Genetics for Reproduction, National Institute for Basic Biology, Okazaki, Japan, Department of Basic Biology, The Graduate University for Advanced Studies (SOKENDAI), Okazaki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MT. Performed the experiments: SN IW TN AT YT. Analyzed the data: SN MT. Contributed reagents/materials/analysis tools: AT YT. Wrote the paper: SN MT.