About the Authors

Susan M. Varnum

Affiliation Department of Cell Biology and Biochemistry, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Bobbie-Jo M. Webb-Robertson

Affiliation Department of Computational Biology and Bioinformatics, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Nancy A. Hessol

Affiliation Department of Clinical Pharmacy, University of California at San Francisco, San Francisco, California, United States of America

Richard D. Smith

Affiliation Department of Biological Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Richard C. Zangar

richard.zangar@pnnl.gov

Affiliation Department of Cell Biology and Biochemistry, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SMV NAH RCZ RDS. Performed the experiments: SMV RCZ. Analyzed the data: SMV B-JMW-R RCZ. Contributed reagents/materials/analysis tools: NAH. Wrote the paper: SMV NAH B-JMW-R RCZ.