About the Authors

Koumei Shirasuna

Affiliation Graduate School of Animal and Food Hygiene, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Japan

Sineenard Jiemtaweeboon

Affiliation Graduate School of Animal and Food Hygiene, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Japan

Sybille Raddatz

Affiliation Clinic for Cattle, University of Veterinary Medicine Hannover, Hannover, Germany

Akane Nitta

Affiliation Graduate School of Animal and Food Hygiene, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Japan

Hans-Joachim Schuberth

Affiliation Clinic for Cattle, University of Veterinary Medicine Hannover, Hannover, Germany

Heinrich Bollwein

Affiliation Institute of Immunology, University of Veterinary Medicine Hannover, Hannover, Germany

Takashi Shimizu

Affiliation Graduate School of Animal and Food Hygiene, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Japan

Akio Miyamoto

akiomiya@obihiro.ac.jp

Affiliation Graduate School of Animal and Food Hygiene, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KS AM. Performed the experiments: KS SJ SR AN. Analyzed the data: KS SJ SR AN. Contributed reagents/materials/analysis tools: HJS HB TS. Wrote the paper: KS.