About the Authors

Dawn L. Cooper

Affiliation NIHR-Leeds Musculoskeletal Biomedical Research Unit, University of Leeds, Leeds, United Kingdom

Stephen G. Martin

Affiliation NIHR-Leeds Musculoskeletal Biomedical Research Unit, University of Leeds, Leeds, United Kingdom

James I. Robinson

Affiliation NIHR-Leeds Musculoskeletal Biomedical Research Unit, University of Leeds, Leeds, United Kingdom

Sarah L. Mackie

Affiliation NIHR-Leeds Musculoskeletal Biomedical Research Unit, University of Leeds, Leeds, United Kingdom

Christopher J. Charles

Affiliation NIHR-Leeds Musculoskeletal Biomedical Research Unit, University of Leeds, Leeds, United Kingdom

Jackie Nam

Affiliation NIHR-Leeds Musculoskeletal Biomedical Research Unit, University of Leeds, Leeds, United Kingdom

YEAR Consortium

Affiliation YEAR Consortium

John D. Isaacs

Affiliation Institute of Cellular Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

Paul Emery

Affiliation NIHR-Leeds Musculoskeletal Biomedical Research Unit, University of Leeds, Leeds, United Kingdom

Ann W. Morgan

a.w.morgan@leeds.ac.uk

Affiliation NIHR-Leeds Musculoskeletal Biomedical Research Unit, University of Leeds, Leeds, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DLC JI PE AWM. Performed the experiments: DLC SGM JIR CC SLM. Analyzed the data: DLC SGM JIR SLM CC JN AWM. Contributed reagents/materials/analysis tools: YEAR JN. Wrote the paper: DLC PE AWM.