About the Authors

Richard L. Mort

Affiliations Division of Reproductive and Developmental Sciences, University of Edinburgh, Edinburgh, United Kingdom, Medical Research Council Human Genetics Unit, Medical Research Council Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Adam J. Bentley

Affiliation Division of Biomedical and Life Sciences, Lancaster University, Lancaster, United Kingdom

Francis L. Martin

Affiliation Centre for Biophotonics, Lancaster University, Lancaster, United Kingdom

J. Martin Collinson

Affiliation Institute of Medical Sciences, University of Aberdeen, Aberdeen, United Kingdom

Panagiotis Douvaras

Affiliation Division of Reproductive and Developmental Sciences, University of Edinburgh, Edinburgh, United Kingdom

Robert E. Hill

Affiliation Medical Research Council Human Genetics Unit, Medical Research Council Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Steven D. Morley

Affiliation School of Clinical Sciences & Community Health, University of Edinburgh, Edinburgh, United Kingdom

Nigel J. Fullwood

Affiliation Division of Biomedical and Life Sciences, Lancaster University, Lancaster, United Kingdom

John D. West

John.West@ed.ac.uk

Affiliation Division of Reproductive and Developmental Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JW RM JC NF SM RH. Performed the experiments: RM AB FM JC PD. Analyzed the data: RM JC. Contributed reagents/materials/analysis tools: JW RM NF. Wrote the paper: RM JW.