About the Authors

Jialou Zhu

Contributed equally to this work with: Jialou Zhu, Zhimao Jiang, Fei Gao, Xueda Hu

Affiliations Beijing Genomics Institute at Shenzhen, Shenzhen, China, College of Life Science, Wuhan University, Wuhan, China

Zhimao Jiang

Contributed equally to this work with: Jialou Zhu, Zhimao Jiang, Fei Gao, Xueda Hu

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Fei Gao

Contributed equally to this work with: Jialou Zhu, Zhimao Jiang, Fei Gao, Xueda Hu

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Xueda Hu

Contributed equally to this work with: Jialou Zhu, Zhimao Jiang, Fei Gao, Xueda Hu

Affiliations Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China, Graduate University of Chinese Academy of Sciences, Beijing, China, Beijing Genomics Institute at Shenzhen, Shenzhen, China

Liang Zhou

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Jiahao Chen

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Huijuan Luo

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Jihua Sun

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Song Wu

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Yonghua Han

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Guangliang Yin

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Maoshan Chen

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Zujing Han

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Xianxin Li

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Yi Huang

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Weixing Zhang

Affiliation Department of Urology, the First Affiliated Hospital of Zhengzhou University, Zhengzhou, China

Fangjian Zhou

Affiliation Department of Urology, Sun Yat-Sen University Cancer Center, Guangzhou, China

Tong Chen

Affiliation Department of Urology, Shenzhen People's Hospital, Shenzhen, China

Pingping Fa

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Yong Wang

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Liang Sun

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Huimin Leng

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Fenghao Sun

Affiliations Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China, Shantou University Medical College, Shantou, China

Yuchen Liu

Affiliations Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China, Department of Urology, Shenzhen People's Hospital, Shenzhen, China

Mingzhi Ye

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Huanming Yang

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Zhiming Cai

Affiliations Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China, Department of Urology, The Second People's Hospital of Shenzhen, Shenzhen, China

Yaoting Gui

zhangxq@genomics.org.cn (XZ); guiyaoting2007@yahoo.com.cn (YG)

Affiliation Guangdong and Shenzhen Key Laboratory of Male Reproductive Medicine and Genetics, Institute of Urology, Peking University Shenzhen Hospital, Shenzhen PKU-HKUST Medical Center, Shenzhen, China

Xiuqing Zhang

zhangxq@genomics.org.cn (XZ); guiyaoting2007@yahoo.com.cn (YG)

Affiliation Beijing Genomics Institute at Shenzhen, Shenzhen, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZC FG XH LZ XL HY YG XZ. Performed the experiments: JZ LZ H. Luo PF H. Leng. Analyzed the data: JZ JC GY MC ZH. Contributed reagents/materials/analysis tools: JZ FG XH LZ H. Luo JS GY XL WZ FZ YW LS FS YL ZJ SW Y. Han TC MY. Wrote the paper: JZ ZJ FG XH. Revised the manuscript and gave final approval of the published version: JZ ZJ FG XH LZ Y. Huang HY YG XZ Y. Han.