About the Authors

Li C. Xia

Affiliation Molecular and Computational Biology Program, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America

Jacob A. Cram

Affiliation Marine and Environmental Biology, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America

Ting Chen

Affiliations Molecular and Computational Biology Program, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America, Tsinghua National Laboratory for Information Science and Technology, Tsinghua University, Beijing, People's Republic of China

Jed A. Fuhrman

Affiliation Marine and Environmental Biology, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America

Fengzhu Sun

fsun@usc.edu

Affiliations Molecular and Computational Biology Program, Department of Biological Sciences, University of Southern California, Los Angeles, California, United States of America, Tsinghua National Laboratory for Information Science and Technology, Tsinghua University, Beijing, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Designed the study: LCX TC JAF FS. Implemented the model and performed the experiments: LCX. Analyzed the data sets: LCX JAC.. Wrote the paper: LCX JAC TC JAF FS.