About the Authors

Emma L. Snary

emma.Snary@ahvla.gsi.gov.uk

Affiliation Centre for Epidemiology and Risk Analysis, Animal Health and Veterinary Laboratories Agency, Addlestone, Surrey, United Kingdom

Vick Ramnial

Affiliation Centre for Epidemiology and Risk Analysis, Animal Health and Veterinary Laboratories Agency, Addlestone, Surrey, United Kingdom

Andrew C. Breed

Affiliation Centre for Epidemiology and Risk Analysis, Animal Health and Veterinary Laboratories Agency, Addlestone, Surrey, United Kingdom

Ben Stephenson

Affiliation Centre for Epidemiology and Risk Analysis, Animal Health and Veterinary Laboratories Agency, Addlestone, Surrey, United Kingdom

Hume E. Field

Affiliation Queensland Centre for Emerging Infectious Diseases, Biosecurity Queensland, Brisbane, Australia

Anthony R. Fooks

Affiliations Wildlife Zoonoses and Vector-borne Diseases Research Group, Department of Virology, Animal Health and Veterinary Laboratories Agency, Addlestone, Surrey, United Kingdom, National Consortium for Zoonosis Research, University of Liverpool, Leahurst, Chester High Road, Neston, United Kingdom

Competing Interests

ARF is an Academic Editor for PLoS ONE.

Author Contributions

Conceived and designed the experiments: ELS ACB VR ARF. Analyzed the data: ELS VR ACB BS . Wrote the paper: ELS VR ACB BS HEF ARF.