About the Authors

Zhongyuan Zhang

Contributed equally to this work with: Zhongyuan Zhang, Kiichiro Teruya

Affiliation Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, Japan

Kiichiro Teruya

Contributed equally to this work with: Zhongyuan Zhang, Kiichiro Teruya

kteruya@grt.kyushu-u.ac.jp

Affiliations Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, Japan, Faculty of Agriculture, Kyushu University, Fukuoka, Japan

Hiroshi Eto

Affiliation Daiichi Sangyo Co., Ltd., Osaka, Japan

Sanetaka Shirahata

Affiliations Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, Japan, Faculty of Agriculture, Kyushu University, Fukuoka, Japan

Competing Interests

This study was supported by Daiichi Sangyo Co., Ltd. and Hiroshi Eto is an employee of Daiichi Sangyo Co., Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: ZZ KT SS. Performed the experiments: ZZ KT. Analyzed the data: ZZ KT. Contributed reagents/materials/analysis tools: ZZ KT HE. Wrote the paper: ZZ KT.