About the Authors

Sarah J. Christianson

Current address: Washington, D.C., United States of America

Affiliation Department of Biology, University of Maryland, College Park, Maryland, United States of America

Cara L. Brand

Current address: Department of Biology, University of Rochester, Rochester, New York, United States of America

Affiliation Department of Biology, University of Maryland, College Park, Maryland, United States of America

Gerald S. Wilkinson

wilkinso@umd.edu

Affiliation Department of Biology, University of Maryland, College Park, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJC GSW. Performed the experiments: SJC CLB. Analyzed the data: SJC GSW. Contributed reagents/materials/analysis tools: GSW. Wrote the paper: SJC GSW.