About the Authors

Stefan W. Metz

Contributed equally to this work with: Stefan W. Metz, Femke Feenstra

Affiliation Laboratory of Virology, Wageningen University, Wageningen, The Netherlands

Femke Feenstra

Contributed equally to this work with: Stefan W. Metz, Femke Feenstra

Affiliation Laboratory of Virology, Wageningen University, Wageningen, The Netherlands

Stephane Villoing

Affiliation Intervet Norbio, Bergen, Norway

Marielle C. van Hulten

Affiliation Intervet International BV, Boxmeer, The Netherlands

Jan W. van Lent

Affiliation Laboratory of Virology, Wageningen University, Wageningen, The Netherlands

Joseph Koumans

Affiliation Intervet International BV, Boxmeer, The Netherlands

Just M. Vlak

Affiliation Laboratory of Virology, Wageningen University, Wageningen, The Netherlands

Gorben P. Pijlman

gorben.pijlman@wur.nl

Affiliation Laboratory of Virology, Wageningen University, Wageningen, The Netherlands

Competing Interests

SV, MH, and JK are employees of Intervet. JV is an employee of Wageningen University and a consultant for Intervet. All experimental work was carried out at Wageningen University, the Netherlands. This did not alter the authors' adherence to all PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GP MH. Performed the experiments: FF SM JL. Analyzed the data: SM FF GP. Contributed reagents/materials/analysis tools: SV JK. Wrote the paper: SM GP JV.