About the Authors

Li Zhu

Contributed equally to this work with: Li Zhu, Wei Hu, Datao Liu

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Biotechnology, Beijing, China

Wei Hu

Contributed equally to this work with: Li Zhu, Wei Hu, Datao Liu

Affiliations State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Biotechnology, Beijing, China, National Institute for the Control of Pharmaceutical and Biological Products, Beijing, China

Datao Liu

Contributed equally to this work with: Li Zhu, Wei Hu, Datao Liu

Affiliation Sine Pharmaceutical Corp. Ltd., Shanghai, China

Wanhong Tian

Affiliation National Institute for the Control of Pharmaceutical and Biological Products, Beijing, China

Gang Yu

Affiliation National Institute for the Control of Pharmaceutical and Biological Products, Beijing, China

Xiankai Liu

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Biotechnology, Beijing, China

Jie Wang

Affiliation National Center of Biomedical Analysis, Beijing, China

Erling Feng

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Biotechnology, Beijing, China

Xuemin Zhang

Affiliation National Center of Biomedical Analysis, Beijing, China

Bei Chen

Affiliation Sine Pharmaceutical Corp. Ltd., Shanghai, China

Ming Zeng

zengming@nicpbp.org.cn (MZ); wanghl@nic.bmi.ac.cn (HW)

Affiliation National Institute for the Control of Pharmaceutical and Biological Products, Beijing, China

Hengliang Wang

zengming@nicpbp.org.cn (MZ); wanghl@nic.bmi.ac.cn (HW)

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Biotechnology, Beijing, China

Competing Interests

DL and BC are employed by Sine Pharmaceutical Corp. Ltd. These relationships do not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HW MZ BC XZ. Performed the experiments: LZ WH DL JW. Analyzed the data: LZ WT XL. Contributed reagents/materials/analysis tools: DL GY BC. Wrote the paper: LZ HW MZ EF.