About the Authors

Go Okada

goookada@hiroshima-u.ac.jp

Affiliations Department of Psychiatry and Neurosciences, Division of Frontier Medical Science, Programs for Biomedical Research, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Corporation, Kawaguchi, Japan

Yasumasa Okamoto

Affiliations Department of Psychiatry and Neurosciences, Division of Frontier Medical Science, Programs for Biomedical Research, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Corporation, Kawaguchi, Japan

Yoshihiko Kunisato

Affiliations Department of Psychiatry and Neurosciences, Division of Frontier Medical Science, Programs for Biomedical Research, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan, Research Fellow of the Japan Society for the Promotion of Science, Tokyo, Japan

Shiori Aoyama

Affiliation Department of Psychiatry and Neurosciences, Division of Frontier Medical Science, Programs for Biomedical Research, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Yoshiko Nishiyama

Affiliation Department of Psychiatry and Neurosciences, Division of Frontier Medical Science, Programs for Biomedical Research, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Shinpei Yoshimura

Affiliation Department of Psychiatry and Neurosciences, Division of Frontier Medical Science, Programs for Biomedical Research, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Keiichi Onoda

Affiliation Department of Neurology, Shimane University, Izumo, Japan

Shigeru Toki

Affiliation Department of Psychiatry and Neurosciences, Division of Frontier Medical Science, Programs for Biomedical Research, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Hidehisa Yamashita

Affiliation Department of Psychiatry and Neurosciences, Division of Frontier Medical Science, Programs for Biomedical Research, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Shigeto Yamawaki

Affiliation Department of Psychiatry and Neurosciences, Division of Frontier Medical Science, Programs for Biomedical Research, Graduate School of Biomedical Sciences, Hiroshima University, Hiroshima, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GO YO YK S. Yoshimura KO ST HY S. Yamawaki. Performed the experiments: GO YK SA YN. Analyzed the data: GO YK. Wrote the paper: GO YO YK KO S. Yoshimura S. Yamawaki.