About the Authors

Benjamin R. Wilcox

Affiliations Southeastern Cooperative Wildlife Disease Study, Department of Population Health, College of Veterinary Medicine, The University of Georgia, Athens, Georgia, United States of America, Daniel B. Warnell School of Forestry and Natural Resources, The University of Georgia, Athens, Georgia, United States of America

Gregory A. Knutsen

Affiliation Agassiz National Wildlife Refuge, U.S. Fish and Wildlife Service, Middle River, Minnesota, United States of America

James Berdeen

Affiliation Minnesota Department of Natural Resources, Wetland Wildlife Populations and Research Group, Bemidji, Minnesota, United States of America

Virginia Goekjian

Affiliation Southeastern Cooperative Wildlife Disease Study, Department of Population Health, College of Veterinary Medicine, The University of Georgia, Athens, Georgia, United States of America

Rebecca Poulson

Affiliation Southeastern Cooperative Wildlife Disease Study, Department of Population Health, College of Veterinary Medicine, The University of Georgia, Athens, Georgia, United States of America

Sagar Goyal

Affiliation Department of Veterinary Population Medicine, University of Minnesota, Saint Paul, Minnesota, United States of America

Srinand Sreevatsan

Affiliations Department of Veterinary Population Medicine, University of Minnesota, Saint Paul, Minnesota, United States of America, Department of Veterinary Biomedical Sciences, University of Minnesota, Saint Paul, Minnesota, United States of America

Carol Cardona

Affiliation Department of Veterinary Biomedical Sciences, University of Minnesota, Saint Paul, Minnesota, United States of America

Roy D. Berghaus

Affiliation Department of Population Health, College of Veterinary Medicine, The University of Georgia, Athens, Georgia, United States of America

David E. Swayne

Affiliation Southeast Poultry Research Laboratory, Agricultural Research Service, U.S. Department of Agriculture, Athens, Georgia, United States of America

Michael J. Yabsley

myabsley@uga.edu

Affiliations Southeastern Cooperative Wildlife Disease Study, Department of Population Health, College of Veterinary Medicine, The University of Georgia, Athens, Georgia, United States of America, Daniel B. Warnell School of Forestry and Natural Resources, The University of Georgia, Athens, Georgia, United States of America

David E. Stallknecht

Affiliation Southeastern Cooperative Wildlife Disease Study, Department of Population Health, College of Veterinary Medicine, The University of Georgia, Athens, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: D. Stallknecht D. Swayne. Analyzed the data: BRW VG RP SG SS CC RDB. Wrote the paper: BRW MJY D. Stallknecht. Collection of field samples: BRW GAK JB.