About the Authors

Paul Lin

pxl57@case.edu

Affiliations Department of Biomedical Engineering, Case Western Reserve University, Cleveland, Ohio, United States of America, Skeletal Research Center, Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America

Diego Correa

Affiliation Skeletal Research Center, Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America

Yuan Lin

Affiliation Stanford Cancer Center, Stanford University, Stanford, California, United States of America

Arnold I. Caplan

Affiliation Skeletal Research Center, Department of Biology, Case Western Reserve University, Cleveland, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PL DC YL AIC. Performed the experiments: PL. Analyzed the data: PL DC YL AIC. Contributed reagents/materials/analysis tools: YL. Wrote the paper: PL.