About the Authors

Helen R. Fryer

Affiliation The Institute for Emerging Infections, Oxford Martin School, Department of Zoology, Oxford University, Oxford, United Kingdom

Angela R. McLean

angela.mclean@zoo.ox.ac.uk

Affiliation The Institute for Emerging Infections, Oxford Martin School, Department of Zoology, Oxford University, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ARM. Analyzed the data: HRF. Wrote the paper: HRF ARM. Performed the mathematical modelling and analysis: HRF.