About the Authors

Frédéric Gazeau

f.gazeau@obs-vlfr.fr

Affiliations Centre National de la Recherche Scientifique-Institut National des Sciences de l'Univers, Laboratoire d'Océanographie de Villefranche, Villefranche-sur-Mer, France, Université Pierre et Marie Curie-Paris 6, Observatoire Océanologique de Villefranche, Villefranche-sur-Mer, France

Jean-Pierre Gattuso

Affiliations Centre National de la Recherche Scientifique-Institut National des Sciences de l'Univers, Laboratoire d'Océanographie de Villefranche, Villefranche-sur-Mer, France, Université Pierre et Marie Curie-Paris 6, Observatoire Océanologique de Villefranche, Villefranche-sur-Mer, France

Mervyn Greaves

Affiliation Department of Earth Sciences, University of Cambridge, Cambridge, England

Henry Elderfield

Affiliation Department of Earth Sciences, University of Cambridge, Cambridge, England

Jan Peene

Affiliation Centre for Estuarine and Marine Ecology, Netherlands Institute of Ecology, Yerseke, The Netherlands

Carlo H. R. Heip

Affiliations Centre for Estuarine and Marine Ecology, Netherlands Institute of Ecology, Yerseke, The Netherlands, Department of Marine Organic Biogeochemistry, Royal Netherlands Institute for Sea Research, Den Burg, The Netherlands

Jack J. Middelburg

Affiliations Centre for Estuarine and Marine Ecology, Netherlands Institute of Ecology, Yerseke, The Netherlands, Faculty of Geosciences, Utrecht University, Utrecht, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FG J-PG JP CHRH JJM. Performed the experiments: FG. Analyzed the data: FG J-PG MG HE CHRH JJM. Contributed reagents/materials/analysis tools: FG J-PG MG HE JP CHRH JJM. Wrote the paper: FG J-PG MG JJM.