About the Authors

Chi-Hun Park

Affiliations Sooam Biotech Research Foundation, Seoul, Republic of Korea, National Institute of Animal Science, RDA, Suwon, Republic of Korea, Department of Agricultural Biotechnology, Research Institute for Agriculture and Life Science, Seoul National University, Seoul, Republic of Korea

Kyung-Jun Uh

Affiliations Sooam Biotech Research Foundation, Seoul, Republic of Korea, Department of Agricultural Biotechnology, Research Institute for Agriculture and Life Science, Seoul National University, Seoul, Republic of Korea

Brendan P. Mulligan

Affiliation Department of Agricultural Biotechnology, Research Institute for Agriculture and Life Science, Seoul National University, Seoul, Republic of Korea

Eui-Bae Jeung

Affiliation College of Veterinary Medicine, Chungbuk National University, Cheongju, Chungbuk, Republic of Korea

Sang-Hwan Hyun

Affiliation College of Veterinary Medicine, Chungbuk National University, Cheongju, Chungbuk, Republic of Korea

Taeyoung Shin

Affiliation Sooam Biotech Research Foundation, Seoul, Republic of Korea

Hakhyun Ka

Affiliation Division of Biological Science and Technology, and Institute of Biomaterials, Yonsei University, Wonju, Republic of Korea

Chang-Kyu Lee

leeck@snu.ac.kr

Affiliation Department of Agricultural Biotechnology, Research Institute for Agriculture and Life Science, Seoul National University, Seoul, Republic of Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: C-HP C-KL. Performed the experiments: C-HP C-KL K-JU. Analyzed the data: E-BJ SH-H TS HK. Contributed reagents/materials/analysis tools: C-HP. Wrote the paper: C-HP BPM C-KL.