About the Authors

Yoko Okamoto

Contributed equally to this work with: Yoko Okamoto, Yoshitomo Shirakashi

Affiliation Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Yoshitomo Shirakashi

Contributed equally to this work with: Yoko Okamoto, Yoshitomo Shirakashi

Affiliation Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Masafumi Ihara

ihara@kuhp.kyoto-u.ac.jp

Affiliation Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Makoto Urushitani

Affiliation Molecular Neuroscience Research Center, Shiga University of Medical Science, Shiga, Japan

Miki Oono

Affiliations Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan, Molecular Neuroscience Research Center, Shiga University of Medical Science, Shiga, Japan

Yasuhiro Kawamoto

Affiliation Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Hirofumi Yamashita

Affiliation Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Shun Shimohama

Affiliation Department of Neurology, Sapporo Medical University School of Medicine, Hokkaido, Japan

Shinsuke Kato

Affiliation Department of Neuropathology, Institute of Neurological Sciences, Faculty of Medicine, Tottori University, Tottori, Japan

Asao Hirano

Affiliation Division of Neuropathology, Department of Pathology, Montefiore Medical Center, New York, New York, United States of America

Hidekazu Tomimoto

Affiliation Department of Neurology, Mie University Graduate School of Medicine, Mie, Japan

Hidefumi Ito

Affiliation Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Ryosuke Takahashi

Affiliation Department of Neurology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MI HT HI SS RT. Performed the experiments: YO YS YK UM MO. Analyzed the data: YO MI. Contributed reagents/materials/analysis tools: SK HY AH. Wrote the paper: YO MI.