About the Authors

Jinxue Long

Current address: Department of Swine Infectious Diseases, Shanghai Veterinary Research Institute, Shanghai, China

Affiliation Biological Mimetics, Inc., Frederick, Maryland, United States of America

Ruth V. Bushnell

Affiliation Biological Mimetics, Inc., Frederick, Maryland, United States of America

John K. Tobin

Affiliation Biological Mimetics, Inc., Frederick, Maryland, United States of America

Keyao Pan

Affiliation Department of Bioengineering and Physics and Astronomy, Rice University, Houston, Texas, United States of America

Michael W. Deem

Affiliation Department of Bioengineering and Physics and Astronomy, Rice University, Houston, Texas, United States of America

Peter L. Nara

Affiliations Biological Mimetics, Inc., Frederick, Maryland, United States of America, Department of Biological Sciences, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Gregory J. Tobin

tobin@bmi-md.com

Affiliations Biological Mimetics, Inc., Frederick, Maryland, United States of America, Department of Biological Sciences, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Competing Interests

RVB, JKT, PLN, and GJT are employees of Biological Mimetics, Inc., a vaccine design strategy corporation. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JL MWD PLN GJT. Performed the experiments: JL RVB JKT KP GJT. Analyzed the data: JL RVB JKT KP MWD PLN GJT. Contributed reagents/materials/analysis tools: JL RVB JKT KP MWD PLN GJT. Wrote the paper: JL RVB JKT KP MWD PLN GJT.