About the Authors
- Sarah E. Belisle
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Contributed equally to this work with: Sarah E. Belisle, Jiangmei Yin
Affiliation Department of Microbiology, University of Washington, Seattle, Washington, United States of America
- Jiangmei Yin
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Contributed equally to this work with: Sarah E. Belisle, Jiangmei Yin
Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America
- Devon J. Shedlock
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Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America
- Anlan Dai
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Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America
- Jian Yan
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Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America
- Lauren Hirao
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Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America
- Michele A. Kutzler
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Affiliation Department of Medicine, Drexel University College of Medicine, Philadelphia, Pennsylvania, United States of America
- Mark G. Lewis
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Affiliation Research Section, Bioqual, Rockville, Maryland, United States of America
- Hanne Andersen
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Affiliation Research Section, Bioqual, Rockville, Maryland, United States of America
- Simon M. Lank
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Affiliation Wisconsin National Primate Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America
- Julie A. Karl
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Affiliation Wisconsin National Primate Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America
- David H. O'Connor
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Affiliation Wisconsin National Primate Research Center, University of Wisconsin-Madison, Madison, Wisconsin, United States of America
- Amir Khan
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Affiliation Inovio Pharmaceuticals, Blue Bell, Pennsylvania, United States of America
- Niranjan Sardesai
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Affiliation Inovio Pharmaceuticals, Blue Bell, Pennsylvania, United States of America
- Jean Chang
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Affiliation Department of Microbiology, University of Washington, Seattle, Washington, United States of America
- Lauri Aicher
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Affiliation Department of Microbiology, University of Washington, Seattle, Washington, United States of America
- Robert E. Palermo
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Affiliation Department of Microbiology, University of Washington, Seattle, Washington, United States of America
- David B. Weiner
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Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America
- Michael G. Katze
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Affiliation Department of Microbiology, University of Washington, Seattle, Washington, United States of America
- Jean Boyer
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* E-mail: boyerj@mail.med.upenn.edu
Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America
Competing Interests
Mark G. Lewis and Hanne Anderson are employed by Bioqual, and Amir Khan and Niranjan Sardesai are employed by Inovio. Inovio provided DNA for the immunization. Primate studies were completed at Bioqual. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.
Author Contributions
Conceived and designed the experiments: JB DBW. Performed the experiments: SEB J. Yin DS AD J. Yan LH MGL LA HA. Analyzed the data: REP MGK JB. Contributed reagents/materials/analysis tools: DBW AK NS MAK. Wrote the paper: JB SEB DS AK NS. Provided guidance: DHO. Completed analysis: JAK JC SML. Completed animal work: HA.