About the Authors

Joram D. Mul

Current address: Department of Medicine, Metabolic Diseases Unit, University of Cincinnati, Cincinnati, Ohio, United States of America

Affiliation Hubrecht Institute, Royal Netherlands Academy of Arts and Sciences and University Medical Center Utrecht, Utrecht, The Netherlands

Susanne E. la Fleur

Current address: Department of Endocrinology and Metabolism, Academic Medical Center, University of Amsterdam, Amsterdam, The Netherlands

Affiliation Department of Neuroscience and Pharmacology, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, The Netherlands

Pim W. Toonen

Affiliation Hubrecht Institute, Royal Netherlands Academy of Arts and Sciences and University Medical Center Utrecht, Utrecht, The Netherlands

Anthonieke Afrasiab-Middelman

Affiliation Department of Cognitive Neuroscience, Donders Institute for Brain, Cognition, and Behavior, Radboud University, Nijmegen, The Netherlands

Rob Binnekade

Affiliation Department of Anatomy and Neurosciences, Center for Neurogenomics and Cognitive Research, Free University Medical Center, Amsterdam, The Netherlands

Dustin Schetters

Affiliation Department of Anatomy and Neurosciences, Center for Neurogenomics and Cognitive Research, Free University Medical Center, Amsterdam, The Netherlands

Michel M. M. Verheij

Affiliation Department of Molecular Animal Physiology, Donders Institute for Brain, Cognition, and Behavior, Radboud University, Nijmegen, The Netherlands

Robert M. Sears

Current address: Center for Neural Science, New York University, New York, New York, United States of America

Affiliation Department of Psychiatry, Ribicoff Research Facilities, Yale University School of Medicine, New Haven, Connecticut, United States of America

Judith R. Homberg

Affiliation Department of Cognitive Neuroscience, Donders Institute for Brain, Cognition, and Behavior, Radboud University, Nijmegen, The Netherlands

Anton N. M. Schoffelmeer

Affiliation Department of Anatomy and Neurosciences, Center for Neurogenomics and Cognitive Research, Free University Medical Center, Amsterdam, The Netherlands

Roger A. H. Adan

Affiliation Department of Neuroscience and Pharmacology, Rudolf Magnus Institute of Neuroscience, University Medical Center Utrecht, Utrecht, The Netherlands

Ralph J. DiLeone

Affiliations Department of Psychiatry, Ribicoff Research Facilities, Yale University School of Medicine, New Haven, Connecticut, United States of America, Department of Neurobiology, Yale University School of Medicine, New Haven, Connecticut, United States of America

Taco J. De Vries

Affiliation Department of Anatomy and Neurosciences, Center for Neurogenomics and Cognitive Research, Free University Medical Center, Amsterdam, The Netherlands

Edwin Cuppen

e.cuppen@hubrecht.eu

Affiliations Hubrecht Institute, Royal Netherlands Academy of Arts and Sciences and University Medical Center Utrecht, Utrecht, The Netherlands, Department of Medical Genetics, University Medical Center Utrecht, Utrecht, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JDM SELF MMMV RMS JRH ANMS RAHA RJD TJDV EC. Performed the experiments: JDM SELF PWT AA-M RB DS MMMV RMS. Analyzed the data: JDM SELF DS MMMV RMS. Contributed reagents/materials/analysis tools: JRH ANMS RAHA RJD TJDV EC. Wrote the paper: JDM SELF MMMV RMS JRH TJDV EC.