About the Authors

D. D. Georgette Lagendijk

georgette.lagendijk@gmail.com

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, University of KwaZulu-Natal, Durban, South Africa

Robin L. Mackey

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, University of KwaZulu-Natal, Durban, South Africa

Bruce R. Page

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, University of KwaZulu-Natal, Durban, South Africa

Rob Slotow

Affiliation Amarula Elephant Research Programme, School of Biological and Conservation Sciences, University of KwaZulu-Natal, Durban, South Africa

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GL BP RS. Performed the experiments: GL. Analyzed the data: GL RM. Wrote the manuscript: GL RM BP RS.