About the Authors

Rhea N. Coler

Contributed equally to this work with: Rhea N. Coler, Sylvie Bertholet

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Sylvie Bertholet

Contributed equally to this work with: Rhea N. Coler, Sylvie Bertholet

Current address: Vaccines and Diagnostics, Novartis, Siena, Italy

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Magdalini Moutaftsi

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Jeff A. Guderian

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Hillarie Plessner Windish

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Susan L. Baldwin

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Elsa M. Laughlin

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Malcolm S. Duthie

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Christopher B. Fox

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Darrick Carter

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Martin Friede

Affiliation Initiative for Vaccine Research, World Health Organization, Geneva, Switzerland

Thomas S. Vedvick

Affiliation Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America

Steven G. Reed

sreed@idri.org

Affiliations Preclinical Biology, Infectious Disease Research Institute, Seattle, Washington, United States of America, Immune Design Corp., Seattle, Washington, United States of America

Competing Interests

Dr. Reed is a founder of, and holds an equity interest in, Immune Design Corp., a licensee of certain rights associated with GLA. This does not alter the authors' adherance to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RNC SB MM SLB HPW. Performed the experiments: MSD MM JAG HPW SB EML CBF TSV. Analyzed the data: RNC MM SB. Wrote the paper: RNC MM SB. Edited the manuscript: SGR DC MF.