About the Authors

Thomas W. Lewis

tom.lewis@aht.org.uk

Affiliation Kennel Club Genetics Centre at the Animal Health Trust, Animal Health Trust, Newmarket, United Kingdom

John A. Woolliams

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Roslin, United Kingdom

Sarah C. Blott

Affiliation Kennel Club Genetics Centre at the Animal Health Trust, Animal Health Trust, Newmarket, United Kingdom

Competing Interests

Hip score and pedigree data was collated and provided by the Kennel Club.

Author Contributions

Conceived and designed the experiments: JAW SCB. Performed the experiments: TWL. Analyzed the data: TWL. Contributed reagents/materials/analysis tools: JAW SCB. Wrote the paper: TWL JAW.