About the Authors

Andrew Bottley

Affiliation School of Pharmacy, University of Nottingham, Nottingham, United Kingdom

Nicola M. Phillips

Affiliation School of Pharmacy, University of Nottingham, Nottingham, United Kingdom

Thomas E. Webb

Affiliation School of Pharmacy, University of Nottingham, Nottingham, United Kingdom

Anne E. Willis

Affiliation School of Pharmacy, University of Nottingham, Nottingham, United Kingdom

Keith A. Spriggs

keith.spriggs@nottingham.ac.uk

Affiliation School of Pharmacy, University of Nottingham, Nottingham, United Kingdom

Competing Interests

Patent application filed by the University of Nottingham. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: AB KAS. Performed the experiments: AB NMP TEW. Analyzed the data: AB NMP TEW KAS. Contributed reagents/materials/analysis tools: AEW KAS. Wrote the paper: KAS.