About the Authors

Angela R. Kamps

Affiliation Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Margaret M. Pruitt

Affiliation Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

John C. Herriges

Current address: Department of Genetics, University of Wisconsin, Madison, Wisconsin, United States of America

Affiliation Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Clark R. Coffman

ccoffman@iastate.edu

Affiliation Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ARK CRC. Performed the experiments: ARK MMP JCH CRC. Analyzed the data: ARK MMP CRC. Contributed reagents/materials/analysis tools: ARK. Wrote the paper: ARK MMP CRC.