About the Authors

Tsuyoshi Hayashi

Contributed equally to this work with: Tsuyoshi Hayashi, Masahiro Nakamichi

Current address: Research Team for Zoonotic Diseases, National Institute of Animal Health (NIAH), National Agriculture and Food Research Organization, Kannondai, Tsukuba, Ibaraki, Japan

Affiliation Laboratory of Microbiology and Immunology, School of Pharmaceutical Sciences, University of Shizuoka, Shizuoka, Japan

Masahiro Nakamichi

Contributed equally to this work with: Tsuyoshi Hayashi, Masahiro Nakamichi

Affiliation Laboratory of Microbiology and Immunology, School of Pharmaceutical Sciences, University of Shizuoka, Shizuoka, Japan

Hirotaka Naitou

Affiliation Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, Shizuoka, Japan

Norio Ohashi

Affiliations Laboratory of Microbiology, Department of Food and Nutritional Sciences, University of Shizuoka, Shizuoka, Japan, Global COE, University of Shizuoka, Shizuoka, Japan

Yasuyuki Imai

Affiliations Laboratory of Microbiology and Immunology, School of Pharmaceutical Sciences, University of Shizuoka, Shizuoka, Japan, Global COE, University of Shizuoka, Shizuoka, Japan

Masaki Miyake

miyakem@ys7.u-shizuoka-ken.ac.jp

Affiliation Laboratory of Microbiology and Immunology, School of Pharmaceutical Sciences, University of Shizuoka, Shizuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MM. Performed the experiments: TH MN HN MM. Analyzed the data: TH MN HN MM. Contributed reagents/materials/analysis tools: NO YI MM. Wrote the paper: TH MM. Critical reading of the manuscript: NO YI.