About the Authors

Behzad Imanian

Contributed equally to this work with: Behzad Imanian, Jean-François Pombert

Affiliation Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Jean-François Pombert

Contributed equally to this work with: Behzad Imanian, Jean-François Pombert

Affiliation Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Patrick J. Keeling

pkeeling@interchange.ubc.ca

Affiliation Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BI JFP PJK. Performed the experiments: BI JFP. Analyzed the data: BI JFP PJK. Wrote the paper: BI JFP PJK.