About the Authors

Chengwei Liang

Contributed equally to this work with: Chengwei Liang, Xiaowen Zhang

Affiliation Qingdao University of Technology and Science, Qingdao, China

Xiaowen Zhang

Contributed equally to this work with: Chengwei Liang, Xiaowen Zhang

Affiliation Yellow Sea Fisheries Research Institute (YSFRI), Chinese Academy of Fishery Sciences, Qingdao, China

Jian Zou

Affiliation Yellow Sea Fisheries Research Institute (YSFRI), Chinese Academy of Fishery Sciences, Qingdao, China

Dong Xu

Affiliation Yellow Sea Fisheries Research Institute (YSFRI), Chinese Academy of Fishery Sciences, Qingdao, China

Feng Su

Affiliation Qingdao University of Technology and Science, Qingdao, China

Naihao Ye

yenh@ysfri.ac.cn

Affiliation Yellow Sea Fisheries Research Institute (YSFRI), Chinese Academy of Fishery Sciences, Qingdao, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CL XZ NY. Performed the experiments: CL XZ NY. Analyzed the data: CL XZ JZ DX FS. Contributed reagents/materials/analysis tools: XZ JZ NY. Wrote the paper: CL XZ.