About the Authors

Ryota Satoh

Contributed equally to this work with: Ryota Satoh, Masafumi Oizumi

Affiliation The University of Tokyo, Kashiwa-shi, Chiba, Japan

Masafumi Oizumi

Contributed equally to this work with: Ryota Satoh, Masafumi Oizumi

Affiliation The University of Tokyo, Kashiwa-shi, Chiba, Japan

Hokto Kazama

Affiliation Department of Neurobiology, Harvard Medical School, Boston, Massachusetts, United States of America

Masato Okada

okada@k.u-tokyo.ac.jp

Affiliations The University of Tokyo, Kashiwa-shi, Chiba, Japan, RIKEN Brain Science Institute, Wako-shi, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: M. Oizumi, H. Kazama, M. Okada. Performed the experiments: R. Satoh. Analyzed the data: R. Satoh. Wrote the paper: R. Satoh, M. Oizumi, H. Kazama, M. Okada.