About the Authors

Daniel P. Smith

Affiliation Molecular and Cellular Biology Program, Oregon State University, Corvallis, Oregon, United States of America

Joshua B. Kitner

Affiliation Department of Microbiology, Oregon State University, Corvallis, Oregon, United States of America

Angela D. Norbeck

Affiliation Biological and Computational Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Therese R. Clauss

Affiliation Biological and Computational Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Mary S. Lipton

Affiliation Biological and Computational Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Michael S. Schwalbach

Affiliation Department of Microbiology, Oregon State University, Corvallis, Oregon, United States of America

Laura Steindler

Affiliation Department of Microbiology, Oregon State University, Corvallis, Oregon, United States of America

Carrie D. Nicora

Affiliation Biological and Computational Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Richard D. Smith

Affiliation Biological and Computational Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Stephen J. Giovannoni

steve.giovannoni@oregonstate.edu

Affiliation Department of Microbiology, Oregon State University, Corvallis, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DPS SJG. Performed the experiments: DPS JBK TRC MSL MSS LS CDN. Analyzed the data: DPS ADN TRC MSL MSS. Contributed reagents/materials/analysis tools: RDS SJG. Wrote the paper: DPS ADN MSS SJG. Developed and made available analysis platforms: RDS.