About the Authors

Angel Gu

Affiliation Department of Immunology, Beckman Research Institute of the City of Hope, Duarte, California, United States of America

Zhifang Zhang

Affiliation Department of Immunology, Beckman Research Institute of the City of Hope, Duarte, California, United States of America

Nan Zhang

Affiliation Department of Immunology, Beckman Research Institute of the City of Hope, Duarte, California, United States of America

Walter Tsark

Affiliation Department of Biology, Beckman Research Institute of the City of Hope, Duarte, California, United States of America

John E. Shively

jshively@coh.org

Affiliation Department of Immunology, Beckman Research Institute of the City of Hope, Duarte, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JS. Performed the experiments: AG ZZ NZ WT. Analyzed the data: AG ZZ. Wrote the paper: JS.