About the Authors

James T. Harper

Affiliations Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada, Department of Biology, Douglas College, New Westminster, British Columbia, Canada

Gillian H. Gile

Affiliation Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Erick R. James

Affiliation Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Kevin J. Carpenter

Affiliation Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Patrick J. Keeling

pkeeling@interchange.ubc.ca

Affiliation Department of Botany, University of British Columbia, Vancouver, British Columbia, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JTH GHG KJC PJK. Performed the experiments: JTH GHG ERJ KJC PJK. Analyzed the data: JTH GHG ERJ KJC PJK. Wrote the paper: PJK.