About the Authors

Bryony W. Soltis

Affiliation Department of Preventive Medicine, Uniformed Services University of Health Sciences, Bethesda, Maryland, United States of America

John W. Sanders

Affiliation Naval Medical Research Center Detachment, Lima, Peru

Shannon D. Putnam

Affiliation U. S. Naval Medical Research Unit No. 2, Jakarta, Indonesia

David R. Tribble

Affiliation Infectious Disease Clinical Research Program, Uniformed Services University of the Health Sciences, Bethesda, Maryland, United States of America

Mark S. Riddle

mark.riddle@med.navy.mil

Affiliation Enteric Diseases Department, Naval Medical Research Center, Silver Spring, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JS SDP DRT. Performed the experiments: JS SDP MSR. Analyzed the data: BWS MSR. Contributed reagents/materials/analysis tools: BWS MSR. Wrote the paper: BWS DRT MSR. Critical editing and revision: JS. Provided critical review and editing: SDP DRT.