About the Authors

Richard Wong

Affiliation Institute of Healthy Ageing, and GEE, University College London, London, United Kingdom

Matthew D. W. Piper

Affiliation Institute of Healthy Ageing, and GEE, University College London, London, United Kingdom

Bregje Wertheim

Affiliations Institute of Healthy Ageing, and GEE, University College London, London, United Kingdom, Evolutionary Genetics, University of Groningen, Groningen, Netherlands

Linda Partridge

l.partridge@ucl.ac.uk

Affiliation Institute of Healthy Ageing, and GEE, University College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RW MDWP LP. Performed the experiments: RW MDWP. Analyzed the data: RW MDWP BW. Contributed reagents/materials/analysis tools: RW MDWP. Wrote the paper: RW.