About the Authors

Timothy R. D. J. Radstake

T.Radstake@reuma.umcn.nl

Affiliations Department of Rheumatology, Radboud University Nijmegen Medical Center, Nijmegen, The Netherlands, The Arthritis Center, Boston University School of Medicine, Boston, Massachusetts, United States of America

Lenny van Bon

Affiliations Department of Rheumatology, Radboud University Nijmegen Medical Center, Nijmegen, The Netherlands, The Arthritis Center, Boston University School of Medicine, Boston, Massachusetts, United States of America

Jasper Broen

Affiliations Department of Rheumatology, Radboud University Nijmegen Medical Center, Nijmegen, The Netherlands, The Arthritis Center, Boston University School of Medicine, Boston, Massachusetts, United States of America

Mark Wenink

Affiliation Department of Rheumatology, Radboud University Nijmegen Medical Center, Nijmegen, The Netherlands

Kim Santegoets

Affiliation Department of Rheumatology, Radboud University Nijmegen Medical Center, Nijmegen, The Netherlands

Yanhui Deng

Affiliation BUMC Flow Cytometry Core Facility, Boston University School of Medicine, Boston, Massachusetts, United States of America

Anila Hussaini

Affiliation The Arthritis Center, Boston University School of Medicine, Boston, Massachusetts, United States of America

Robert Simms

Affiliation The Arthritis Center, Boston University School of Medicine, Boston, Massachusetts, United States of America

William W. Cruikshank

Affiliation The Pulmonary Center, Boston University School of Medicine, Boston, Massachusetts, United States of America

Robert Lafyatis

Affiliation The Arthritis Center, Boston University School of Medicine, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TR LvB JB MW KS WC RL. Performed the experiments: TR LvB JB MW KS YD WC. Analyzed the data: TR LvB YD. Contributed reagents/materials/analysis tools: TR AH RS RL. Wrote the paper: TR LvB JB RS RL.